Afrotropical mayfly families

List of Afrotropical mayfly families, genera and species, with synonyms (including authors and dates for all specific name changes).

 Compiled by Helen Barber-James, Albany Museum, Grahamstown, South Africa.

Updated August 2010

This checklist aims to include all known species found in the Afrotropical region, as defined by Crosskey and White (1977). Species which occur north of this boundary have been excluded. If the taxon is found on one of the Afrotropical islands, this is indicated by [Madagascar], [Mauritius], [Seychelles] or [Comoros]. Otherwise it is found on the African mainland. Several species awaiting description have not been included, but this list will continue to be updated.

Note that in the list below, the current valid names of the species (indicated by ?) are followed by subordinate names when these exist (indicated by ?). The subordinate names are followed by an abbreviation in parentheses, indicating their status. All names, valid or subordinate, are followed by the actual author of that name and the date that name was published. The authors responsible for the latest names changes are indicated at the end of each list of synonyms.

Definitions of abbreviations and terms used:

(comb.) combination: a name recombined with a generic name different from the original;

(dub.) nomen dubium: a name that is of dubious status in that it cannot presently be determined what the species named as such actually is; although all nomina dubia have a generic name associated with them, keep in mind that some of them cannot actually be placed to any genus with confidence due to the lack, or loss, of original material on which the names were based;

(hom.) homonym: the name (as the unique combination of genus and species names) is or becomes the same as one used for another species at an earlier time; the subordinate homonym, therefore, had to be replaced with a new name, see

(renam.), below; the species name that has priority is also indicated within the parentheses along with (hom.);

(id.) misidentification: another valid species (not a subordinate name) with which the species has incorrectly been associated in the past;

(orig.) original: the original name, including the original name in its correct form if it has been changed from an incorrectly formed name;

(renam.) renamed: a new name given because the original or recombination of the original is a secondary homonym (the same as that which has been given to another species previously);

(spell.) spelling: a misspelled or incorrect form of the name;

(stat.) status: represents a change from species to subspecies status within the same species;

(syn.) synonym: another name that was proposed independently for the same species. Normally, the earliest published name takes priority.

(in part): some of the members of this species were originally described as another species.

[sic]: name written as stated by author, even though incorrect

†: extinct


Afrotropical Region
The Afrotropical Region


Suborder PISCIFORMA

Superfamily SIPHLONUROIDEA

Family BAETIDAE

1. GenusAcanthiops Waltz and McCafferty, 1987

? Acanthiops cooperi (Gillies & Wuillot, 1997)

? Platycloeon cooperi Gillies & Wuillot, 1997 (orig.)

? Acanthiops cooperi Lugo-Ortiz & McCafferty, 1998 (comb.)

? Acanthiops delphinae (Gattolliat, 2000) [Madagascar]

? Afroptiloides delphinae Gattolliat, 2000 (orig.)

? Acanthiops delphinae Lugo-Ortiz, Barber-James, McCafferty & de Moor, 2001 (comb.)

? Acanthiops elgonensis Lugo-Ortiz &McCafferty 1998

? Centoptilum sp. no. 3. Demoulin 1964 (syn.)

? Acanthiops elgonensis Lugo-Ortiz & McCafferty 1998 (orig.)

? Afroptiloides elgonensis Gillies 1999 (comb.)

? Acanthiops elgonensis Lugo-Ortiz, Barber-James, McCafferty & de Moor, 2001 (comb.)

? Acanthiops erepens (Gillies, 1990) [South Africa]

? Baetis cataractae Crass, 1947 (in part)

? Baetis sp. A. Kimmins, 1955 (in part)

? Afroptilum erepens Gillies, 1990 (orig.)

? Platycloeon erepens Gillies & Wuillot, 1997 (comb.)

? Acanthiops erepens Lugo-Ortiz & McCafferty, 1998 (comb.)

? Acanthiops faro Barber-James & McCafferty, 2001

? Acanthiops gosei McCafferty, 2002

? Acanthiops griffithsi Lugo-Ortiz &McCafferty 1998

? Afroptiloides griffithsi Gillies 1999 (orig.)

? Acanthiops griffithsi Lugo-Ortiz, Barber-James, McCafferty & de Moor, 2001 (comb.)

? Acanthiops io Lugo-Ortiz & McCafferty, 2001

? Acanthiops marlieri (Demoulin, 1967)

? Centroptilium marlieri Demoulin 1967 (orig.)

? Acanthiops marlieri Waltz & McCafferty, 1987 (comb.)

? Acanthiops namorona (Gattolliat, 2000) [Madagascar]

? Afroptiloides namorona Gattolliat, 2000 (orig.)

? Acathiops namorona Lugo-Ortiz, Barber-James, McCafferty & de Moor, 2001 (comb.)

? Acanthiops spinosum (Gattolliat, 2000) [Madagascar]

? Afroptiloides spinosum Gattolliat, 2000 (orig.)

? Acanthiops spinosum Lugo-Ortiz, Barber-James, McCafferty & de Moor, 2001 (comb.)

? Acanthiops tsitsa Barber-James & McCafferty, 1997 [South Africa]

? Afroptiloides tsita [sic] Gillies, 1999 (spell.)

? Acanthiops tsitsa Lugo-Ortiz, Barber-James, McCafferty & de Moor, 2001 (comb.)

? Acanthiops variegatus (Gillies, 1991)

? Afroptilum variegatum Gillies, 1991 (orig.)

? Acanthiops variegatus Barber-James & McCafferty, 1997 (comb.)

? Afroptiloides variegatum [sic] Gillies, 1999 (comb.)

? Acanthiops variegatus Lugo-Ortiz, Barber-James, McCafferty & de Moor, 2001 (comb.)

? Acanthiops varius (Crass, 1947) [South Africa]

? Centroptilum varium Crass, 1947 (orig.)

? Acentrella sp. Demoulin, 1956 (syn.)

? Afroptilum (Afroptiloides) varium Gillies, 1990 (comb.)

? Acanthiops varius McCafferty & de Moor, 1995 (comb.)

? Acanthiops tsitsa Lugo-Ortiz & McCafferty, 1998 (id.)

? Afroptiloides varium [sic] Gillies, 1999 (comb.)

? Acanthiops varius Lugo-Ortiz, Barber-James, McCafferty & de Moor, 2001 (comb.)

? Acanthiops zomba Lugo-Ortiz & McCafferty, 1998

? Acentrella sp. A. Kimmins, 1955 (syn.)

? Baetis sp. A. Kimmins, 1955 (in part) (syn.)

? Centroptilum sp. A. Demoulin, 1970 (syn.)

? Afroptilum (Afroptiloides) sp. A. Gillies, 1990 (syn.)

? Acanthiops zomba Lugo-Ortiz & McCafferty, 1998 (orig.)

? Afroptiloides njombae [sic.] Gillies, 1999. (comb., spell.)

? Acanthiops zomba Lugo-Ortiz, Barber-James, McCafferty & de Moor, 2001 (comb.)

2. Genus Adnoptilum Gattolliat & Monaghan 2010

? Adnoptilum ottkei Gattolliat and Monaghan 2010

? Adnoptilum thomasorum Gattolliat and Monaghan 2010

3. Genus Afrobaetodes Demoulin, 1970

? Afrobaetodes berneri Demoulin, 1970

? Afrobaetodes pugio Gillies, 1991 (syn.)

? Afrobaetodes intermedius Lugo-Ortiz & McCafferty, 1996 (syn.)

? Afrobaetodes lenae Gattolliat & Sartori, 1999 [Madagascar]

4. GenusAfroptilum Gillies, 1990

? Afroptilum biarcuatum (Kopelke, 1980)

? Centroptilum biarcuatum Kopelke, 1980 (orig.)

? Afroptilum biarcuatum Gillies, 1990 (comb.)

? Afroptilum bicorne (Ulmer, 1909)

? Centroptilum bicorne Ulmer, 1909 (orig.)

? Afroptilum bicorne Gillies, 1990 (comb.)

? Afroptilum boettgeri (Kopelke, 1980)

? Centroptilum boettgeri Kopelke, 1980 (orig.)

? Afroptilum boettgeri Gillies, 1990 (comb.)

? Afroptilum confusum Lugo-Ortiz & McCafferty, 1998 [Madagascar]

? Afroptilum dicentrum (Demoulin, 1956)

? Centroptilum dicentrum Demoulin, 1956 (orig.)

? Afroptilum dicentrum Gillies, 1990 (comb.)

? Afroptilum gilberti Gattolliat & Sartori, 1999 [Madagascar]

? Afroptilum lepidum Lugo-Ortiz & McCafferty, 1998 [Madagascar]

? Afroptilum mathildae Gattolliat & Sartori, 1999 [Madagascar]

? Afroptilum parvum (Crass, 1947) [South Africa]

? Centroptilum parvum Crass, 1947 (orig.)

? Afroptilum parvum Gillies, 1990 (comb.)

? Afroptilum sudafricanum (Lestage, 1924) [South Africa]

? Centroptilum sudafricanum Lestage, 1924 (orig.)

? Centroptilum montanum Kimmins, 1960 (syn.)

? Afroptilum sudafricanum Gillies, 1990 (comb.)

5. Genus Baetis Leach, 1815

? Baetis aeneus Navàs, 1936

? Baetis harrisoni Barnard, 1932 [South Africa]

? Baetis lawrencei Crass, 1947 [South Africa]

? Baetis magae (Barnard, 1932) [South Africa]

? Pseudocloeon magae Barnard, 1932 (orig.)

? Baetis magae Gillies, 1994 (comb.)

? Baetis monikae Kopelke, 1980

? Baetis parvulus Crass, 1947 [South Africa]

? Baetis permultus Kopelke, 1980

? Baetis pseudogemellus Soldàn, 1977

? Baetis spatulatus Gillies, 1994

? Baetis tripunctatus Gillies, 1994

6. Genus Barnumus Lugo-Ortiz & McCafferty, 1998

? Barnumus editus McCafferty & Lugo-Ortiz, 1998 [South Africa]

7. Genus BugilliesiaLugo-Ortiz &McCafferty, 1996

? Bugilliesia biloba Gattolliat, 2006 ?,? imago. Nymph unknown. [Mali, Gunea, Ivory Coast]

? Bugilliesia cavalliensis Gattolliat, 2006 ?,? imago. Nymph unknown. [Ivory Coast, Guinea]

? Bugilliesia grisea (Gillies, 1990) ?,? imago. Nymph unknown. [Tanzania]

? Afroptilum griseum Gillies, 1990 (orig.)

? Bugilliesia grisea Lugo-Ortiz & McCafferty, 1996 (comb.)

? Bugilliesia guineensis (Gillies, 1990) ? imago, nymph. [Guinea, Ivory Coast, Togo]

? Afroptilum guineensis Gillies, 1990 (orig.)

? Bugilliesia guineensis Lugo-Ortiz & McCafferty, 1996 (comb.)

? Bugilliesia margaretae Gattolliat & Barber-James, 2009 [South Africa]

? Bugillesia mirandei (Lugo-Ortiz & McCafferty, 1997) [Madagascar]

? Cheleocloeon mirandei Lugo-Ortiz & McCafferty, 1997 orig.

? Bugillesia mirandei Gattolliat, Barber-James & Monaghan, 2009 (comb)

? Bugillesia nitida (Ulmer, 1916) [Democratic Republic of the Congo].

? Centroptilum nitidum Ulmer, 1916 (orig.)

? Cloeon nigroalbum Navás, 1932 (syn.)

? Cloeon bredoanum Navás, 1933 (syn.)

? Centroptilum nitidum Demoulin, 1957 (comb.)

? Afroptilum nitidum Gillies, 1990 (comb.)

? Bugillesia nitida Lugo-Ortiz & McCafferty, 1996 (comb.)

? Bugilliesia notabilis (Kimmins, 1956) ?,? imago. Nymph unknown.[Uganda, Tanzania, Guinea, Mali, Ivory Coast]

? Centroptilum notabile Kimmins, 1956 (orig.)

? Afroptilum notabile Gillies, 1990 (comb.)

? Bugilliesia notabilis Lugo-Ortiz & McCafferty, 1996 (comb.)

? Bugilliesia sudanensis (Ulmer, 1916) ? imago. Nymph unknown. [Tanzania, Guinea, Gambia, Ivory Coast]

? Centroptilum sudanense Ulmer, 1916 (orig.)

? Afroptilim sudanense Gillies, 1990 (comb.)

? Bugilliesia sudanensis Lugo-Ortiz & McCafferty, 1996 (comb.)

? Bugilliesia truncata Gattolliat, 2006 ?,? imago. Nymph unknown. [Guinea, Sierra Leone].

8. Genus Centroptiloides Lestage, 1918

? Centroptiloides bifasciata (Esben-Petersen, 1913) [South Africa]

? Centroptilum bifasciatum Esben-Petersen, 1913 (orig.)

? Centroptiloides bifasciatus Lestage, 1918 (comb.)

? Centroptiloides bifasciatum Ulmer, 1920 (renam.)

? Haplobaetis umbratus Navás, 1922 (syn.)

? Centroptiloides bifasciata Navás, 1922 (renam.)

? Centroptiloides marginata Lestage, 1924 (syn.)

? Centroptiloides collarti Navás, 1930 (syn.)

? Centroptiloides umbratus Lestage, 1945 (syn.)

? Centroptiloides bifasciatum (form hyalinum) Crass, 1947 (stat.)

? Centroptiloides bifasciata (form marginata) Demoulin, 1957 (stat.)

? Centroptiloides ornatus Gillies, 2001

9. Genus Cheleocloeon Wuillot & Gillies, 1993

? Cheleocloeon carinatum Wuillot, 1993

? Cheleocloeon excisum (Barnard, 1932) [South Africa]

? Centroptilum excisum Barnard, 1932 (orig.)

? Afroptilum excisum Gillies, 1990 (comb.)

? Cheleocloeon excisum Lugo-Ortiz & McCafferty, 1997 (comb.)

? Cheleocloeon falcatum (Crass, 1947) [South Africa]

? Centroptilum falcatum Crass 1947 (orig.)

? Afroptilum falcatum Gillies 1990 (comb.)

? Cheleocloeon falcatum Lugo-Ortiz & McCafferty 1998 (comb.)

? Cheleocloeon littorale McCafferty, 2000

? Cheleocloeon madagascariense Gattolliat & Monaghan, 2009 [Madagascar]

? Cheleocloeon sigiense Gillies 2001

? Cheleocloeon soldáni Gattolliat & Sartori 2008

? Cheleocloeon yolandae Wuillot, 1993

10. Genus Cloeodes Traver,1938

? Cloeodes bicoloratus Gattolliat, 2001 [Madagascar]

? Cloeodes dentatus (Kimmins,1956) [South Africa]

? Cloeon dentatum Kimmins, 1956

? Potamocloeon dentatum Gillies, 1990

? Afroptilum plumosum Wuillot, 1993

? Maliqua plumosa Lugo-Ortiz & McCafferty, 1997

? Maliqua abdallahi (McCafferty, 2000)

? Cloeodes freitagae Gattolliat, 2001 [Madagascar]

? Cloeodes inzingae (Crass, 1947) [South Africa]

? Pseudocloeon inzingae Crass 1947

? Pseudocloeon saxophilum Agnew 1961

? Baetis inzingae Gillies 1994

? Baetis saxophilus Gillies 1994

? Cloeodes inzingae Waltz & McCafferty 1994

? Cloeodes saxophilus de Moor & McCafferty 1996

? Cloeodes portabilis Lugo-Ortiz & McCafferty, 1999 [Madagascar]

? Cloeodes pseudogladius Gattolliat, 2001 [Madagascar]

? Cloeodes pusillum (Navás, 1930) [South Africa]

? Cloeon pusillus Navás, 1930

? Cloeon delicatissimum Barnard, 1932 (syn.)

? Afrobaetodes pusillus Gillies, 1979

? Cloeodes pusillum Jacobus, McCafferty & Gattolliat, 2006

11. Genus CloeonLeach 1815

? Cloeon aeneum Barnard, 1932 [South Africa]

? Cloeon agnewi Hubbard, 1973 [South Africa]

? Austrocloeon exiguum Crass 1947 (nec Navás 1918) (orig.)

? Cloeon exiguum Demoulin 1970 (comb.)

? Cloeon agnewi Hubbard 1973 (hom, renam.)

? Cloeon amaniensis Gillies, 1985

? Cloeon areolatum Navàs, 1930

? Cloeon bellum Navàs, 1931

? Cloeon cambouei Navàs, 1930 [Madagascar] (nomen dubium - Lugo-Ortiz & McCafferty, 1998)

? Cloeon chaplini Barnard, 1932 [South Africa]

? Cloeon crassi Agnew, 1961 [South Africa]

? Austrocloeon sp. A. Allanson, 1961 (syn.)

? Cloeon durani Navás, 1926 [Madagascar]

? Cloeon durani Navás, 1926 (orig.)

? Cloeon waterloti Demoulin, 1966 (syn.)

? Cloeon durani Gattolliat & Rabeantoandro, 2002 (redescribed)

? Cloeon elevatum Agnew, 1961 [South Africa]

? Cloeon emmanueli Lugo-Ortiz & McCafferty, 1998 [Madagascar]

? Cloeon gambiae Gillies, 1980

? Cloeon irretitum Navàs, 1936 [Madagascar] (nomen dubium - Lugo-Ortiz & McCafferty, 1998)

? Cloeon lacunosum Barnard, 1932 [South Africa]

? Cloeon madhouae McCafferty & Mauremootoo, 2000 [Mauritius]

? Cloeon perkinsi Barnard, 1932 [South Africa]

? Cloeon rhodesiae Barnard, 1932 [South Africa]

? Cloeon carneum Navás, 1936 (syn.)

? Cloeon stigmale Navás, 1936 (syn.)

? Cloeon scitulum Kimmins, 1955

? Cloeon smaeleni Lestage, 1924 [Africa & Madagascar]

? Procloeon smaeleni Kimmins, 1960 (orig.)

? Procloeon fraudulentum Demoulin, 1957 (syn.)

? Cloeon affine Navás, 1930 (syn.)

? Cloeon incertum Demoulin, 1957 (syn.)

? Cloeon punctatum Navás, 1931 (syn.)

? Cloeon smaeleni Gattolliat & Rabeantoandro, 2002 (comb.)

? Cloeon tanzaniae Gillies, 1985

? Cloeon virgiliae (Barnard, 1932) [South Africa]

? Austrocloeon virgiliae Barnard, 1932 (orig.)

? Austrocloeon paludinosum Crass, 1947 (syn.)

? Cloeon virgiliae Demoulin, 1970 (comb.)

? Cloeon viridellum Lestage, 1923

12. Genus Crassabwa Lugo-Ortiz & McCafferty, 1996

? Crassabwa badia (Kopelke, 1980)

? Centroptilum badium Kopelke, 1980 (orig.)

? Crassabwa badia Lugo-Ortiz & McCafferty, 1996 (comb.)

? Crassabwa flava (Crass, 1947) [South Africa]

? Centroptilum flavum Crass, 1947 (orig.)

? Afroptilum flavum Gillies, 1990 (comb.)

? Crassabwa flava Lugo-Ortiz & McCafferty, 1996 (comb.)

? Crassabwa loweae (Kimmins, 1949)

? Centroptilum loweae Kimmins, 1949 (orig.)

? Crassabwa loweae Lugo-Ortiz & McCafferty, 1996 (comb.)

? Crassabwa vitrea Navàs, 1930

? Centroptilum vitreum Navàs, 1930 (orig.)

? Crassabwa vitrea Lugo-Ortiz & McCafferty, 1996 (comb.)

13. Genus Dabulamanzia Lugo-Ortiz & McCafferty, 1996

? Dabulamanzia babaora (Wuillot, 1993)

? Afroptilum babaorum Wuillot, 1993 (orig.)

? Dabulamanzia babaora Lugo-Ortiz & McCafferty, 1996 (comb.)

? Dabulamanzia concolorata Gattolliat, 2000 [Madagascar]

? Dabulamanzia fica Lugo-Ortiz & McCafferty, 1996 [South Africa]

? Dabulamanzia gigantea Gattolliat, 2000 [Madagascar]

? Dabulamanzia gladius Gattolliat, 2000 [Madagascar]

? Dabulamanzia helenae Lugo-Ortiz & McCafferty, 1996 [South Africa]

? Dabulamanzia improvida Lugo-Ortiz & McCafferty, 1997 [Madagascar]

?Dabulamanzia duci Gattolliat & Elouard, 1999 [Madagascar]

? Nesydemius polhemusorum Lugo-Ortiz & McCafferty, 1998 [Madagascar]

? Dabulamanzia indusii (Crass, 1947) [South Africa]

? Centroptilum indusii Crass, 1947 (orig.)

? Afroptilum indusii Gillies, 1990 (comb.)

? Dabulamanzia indusii Lugo-Ortiz & McCafferty, 1996 (comb.)

? Dabulamanzia media (Crass, 1947) [South Africa]

? Centroptilum medium Crass, 1947 (orig.)

? Afroptilum medium Gillies, 1990 (comb.)

? Dabulamanzia media Lugo-Ortiz & McCafferty, 1996 (comb.)

? Dabulamanzia tarsale (Gillies, 1990) [South Africa]

? Afroptilum tarsale Gillies, 1990 (orig.)

? Dabulamanzia tarsale Lugo-Ortiz & McCafferty, 1996 (comb.)

14. Genus Delouardus Lugo-Ortiz & McCafferty, 1999 [Madagascar]

? Delouardus djabala Lugo-Ortiz & McCafferty, 1999 [Madagascar]

15. Genus Demoreptus Lugo-Ortiz & McCafferty 1997

? Demoreptus capensis (Barnard, 1932) [South Africa]

? Acentrella capensis Barnard, 1932 (orig.)

? Baetis capensis Demoulin, 1970 (comb.)

? Acentrella capensis McCafferty & de Moor, 1995 (comb.)

? Demoreptus capensis Lugo-Ortiz & McCafferty, 1997 (comb.)

? Demoreptus monticola (Crass, 1947) [South Africa]

? Acentrella monticola Crass, 1947 (orig.)

? Baetis monticola Demoulin, 1970 (comb.)

? Acentrella monticola McCafferty & de Moor, 1995 (comb.)

? Demoreptus monticola Lugo-Ortiz & McCafferty, 1997 (comb.)

? Demoreptus natalensis (Crass, 1947) v

? Acentrella natalensis Crass, 1947 (orig.)

? Baetis natalensis Demoulin, 1970 (comb.)

? Acentrella natalensis McCafferty & de Moor, 1995 (comb.)

? Demoreptus natalensis Lugo-Ortiz & McCafferty, 1997 (comb.)

16. Genus Demoulinia Gillies, 1990

? Demoulinia assimilis Gattolliat (2003) [Madagascar]

? Demoulinia crassi (Demoulin, 1970) [South Africa]

? Centroptilum pulchrum Crass, 1947 (nec Eaton, 1869) (orig.)

? Centroptilum crassi Demoulin, 1970 (hom, renam.)

? Demoulinia crassi Gillies, 1990 (comb.)

? Demoulinia insularis Lugo-Ortiz & McCafferty, 1998 [Madagascar]

17. Genus Dicentroptilum Wuillot & Gillies, 1994

? Dicentroptilum clandestinum Lugo-Ortiz & McCafferty, 2001

? Dicentroptilum decipiens (Gillies, 1990)

? Dicentroptilum imanishii McCafferty, 2002

? Dicentroptilum merina Lugo-Ortiz & McCafferty, 1998 [Madagascar]

? Dicentroptilum papillosum Wuillot, 1994

? Dicentroptilum spinulosum (Demoulin, 1970) [South Africa]

? Centroptiloides spinulosa Demoulin, 1970 (orig.)

? Afroptilum spinulosum Gillies, 1990 (comb.)

? Dicentroptilum spinulosum Wuillot & Gillies, 1994 (comb.)

18. Genus Echinopus Gattolliat (2002) [Madagascar]

? Echinopus giboni Gattolliat (2002) [Madagascar]

? Echinopus minutus Gattolliat (2002)[Madagascar]

19. Genus Edmulmeatus Lugo-Ortiz & McCafferty, 1997 [Madagascar]

? Edmulmeatus grandis Lugo-Ortiz & McCafferty, 1997 [Madagascar]

20. Genus Glossidion Lugo-Ortiz & McCafferty, 1998

? Glossidion demoulini Lugo-Ortiz & McCafferty, 1998

? Glossidion mysticum Lugo-Ortiz & McCafferty, 1998

21. Genus Guloptiloides Gattolliat & Sartori, 2000

? Guloptiloides gargantua Gattolliat & Sartori, 2000 [Madagascar]

22. Genus Herbrossus McCafferty & Lugo-Ortiz, 1998 [Madagascar]

? Herbrossus christinae Gattolliat & Sartori, 1998 [Madagascar]

? Herbrossus edmundsorum McCafferty & Lugo-Ortiz, 1998 [Madagascar]

? Herbrossus elouardi Gattolliat & Sartori, 1998 [Madagascar]

23. Genus Kivuiops McCafferty & Lugo-Ortiz, 1996

? Kivuiops elouardi (Gillies, 1989) ?,? imago. Nymph unknown. [Ivory Coast]

? Rhithrocloeon elouardi Gillies, 1989 (orig.)

? Kivua elouardi McCafferty & Lugo-Ortiz, 1996 (comb.)

? Kivuiops insueta (Kopelke, 1980)

? Cloeon insuetum Kopelke, 1980 (orig.)

? Kivua insueta McCafferty & Lugo-Ortiz, 1996 (comb.)

24. Genus LabiobaetisNovikova & Kluge, 1987

? Labiobaetis dambrensis Gattolliat, 2001 [Madagascar]

? Labiobaetis fabulosus Lugo-Ortiz & McCafferty, 1997 [Madagascar]

? Labiobaetis fabulosus Lugo-Ortiz & McCafferty, 1997 (orig.)

? Pseudocloeon fabulosum Lugo-Ortiz, McCafferty & Waltz, 1999 (comb.)

? Labiobaetis fabulosus Gattolliat, 2001 (comb.)

? Labiobaetis gambiae (Gillies, 1993)

? Baetis gambiae Gillies, 1993 (orig.)

? Labiobaetis gambiae Lugo-Ortiz & McCafferty, 1997 (comb.)

? Labiobaetis gilliesi Gattolliat, 2001 [Madagascar]

? Labiobaetis longicercus Gattolliat, 2001 [Madagascar]

? Labiobaetis nigrocercus Gattolliat, 2001 [Madagascar]

? Labiobaetis plumbago Lugo-Ortiz & McCafferty, 1997 [Madagascar]

? Labiobaetis plumbago Lugo-Ortiz & McCafferty, 1997 (orig.)

? Pseudocloeon plumbago Lugo-Ortiz, McCafferty & Waltz, 1999 (comb.)

? Labiobaetis plumbago Gattolliat, 2001 (comb.)

? Labiobaetis punctatus Gattolliat, 2001 [Madagascar]

? Labiobaetis vulgaris Gattolliat, 2001 [Madagascar]

25. GenusMickiops McCafferty, Lugo-Ortiz & Barber-James, 1997

? Micksiops bicaudatum (Gillies, 1990)

? Afroptilum (Afroptiloides) bicaudatum Gillies, 1990 (orig.)

? Mickiops bicaudatum McCafferty, Lugo-Ortiz & Barber-James, 1997 (comb.)

26. Genus Mutelocloeon Gillies & Elouard, 1990

? Mutelocloeon bihoumi Gillies & Elouard, 1990 ?,? imago, nymph. [Mali, Guinea, Sierra Leone]

? Mutelocloeon corbeti (Kimmins, 1956)

? Centroptilum corbeti Kimmins, 1956 (orig.)

? Mutelocloeon corbeti Gillies & Elouard, 1990 (comb.)

? Mutelocloeon thomasorum Lugo-Ortiz & McCafferty, 1997 [Madagascar] incertae sedis Gattolliat, 2006.

27. Genus Nesoptiloides Demoulin, 1973 [Madagascar]

? Nesoptiloides electroptera (Demoulin, 1966) [Madagascar]

? Centroptilum electropterum Demoulin, 1966 (orig.)

? Nesoptiloides intermedia Demoulin, 1973 (syn.)

? Nesoptiloides electroptera Gattolliat & Sartori, 1999 (comb.)

28. Genus Nigrobaetis Novikova & Kluge, 1987

? Nigrobaetis arabiensis Gattolliat & Sartori, 2008

? Nigrobaetis bethunae Lugo-Ortiz & de Moor, 2000 [South Africa]

? Nigrobaetis colonus Gattolliat, 2004 [La Réunion]

? Nigrobaetis cryptus Gattolliat, 2004 [Madagascar]

? Nigrobaetis harasab (Soldàn, 1977)

? Baetis harasab Soldàn, 1977 (orig.)

? Nigrobaetis harasab Lugo-Ortiz & de Moor, 2000 (comb.)

29. Genus Ophelmatostoma Waltz & McCafferty, 1987

? Ophelmatostoma camerunense (Ulmer, 1920) [South Africa]

? Pseudocloeon camerunense Ulmer, 1920 (orig.)

? Pseudocloeon sp. A. Kimmins, 1955 (syn.)

? Ophelmatostoma kimminsi Waltz & McCafferty, 1987 (syn.)

? Ophelmatostoma camerunense Gillies, Elouard & Wuillot, 1990

30. Genus Peuhlella Lugo-Ortiz & McCafferty, 1998

? Peuhlella christinae (Wuillot, 1993)

? Afroptilum christinae Wuillot, 1993 (orig.)

? Peuhlella christinae Lugo-Ortiz & McCafferty, 1998 (comb.)

31. Genus Procloeon Bengtsson, 1915

? Procloeon africanum (Esben-Petersen, 1913) [South Africa]

? Cloeon africanum Esben-Petersen, 1913 (orig.)

? Cloeon marginale Ulmer, 1916 (syn.)

? Austrocloeon africanum Barnard, 1932 (comb.)

? Cloeon africanum Demoulin, 1970 (comb.)

? Procloeon africanum Gillies, 1997 (comb.)

? Procloeon cylindroculum Kimmins, 1955

? Procloeon sylvicola Gillies, 1997

32. Genus Pseudocloeon Klapàlek, 1905

? Pseudocloeon bellum (Barnard, 1932) [South Africa]

? Baetis bellus Barnard, 1932 (orig.)

? Pseudocloeon bellum Lugo-Ortiz, McCafferty & Waltz, 1999 (comb.)

? Pseudocloeon boussoulium (Gillies, 1993)

? Baetis boussoulius Gillies, 1993 (orig.)

? Labiobaetis boussoulius Lugo-Ortiz & McCafferty, 1997 (comb.)

? Pseudocloeon boussoulium Lugo-Ortiz, McCafferty & Waltz, 1999 (comb.)

? Pseudocloeon cataractae (Crass, 1947) [South Africa]

? Baetis cataractae Crass, 1947 (orig.)

? Acanthiops erepens (in part) Lugo-Ortiz, Barber-James, McCafferty & de Moor, 2001 (id.)

? Pseudocloeon cataractae Lugo-Ortiz & de Moor, 2001 (comb.)

? Pseudocloeon elouardi (Gillies, 1993)

? Baetis elouardi Gillies, 1993 (orig.)

? Labiobaetis elouardi Lugo-Ortiz & McCafferty, 1997 (comb.)

? Pseudocloeon elouardi Lugo-Ortiz, McCafferty & Waltz, 1999 (comb.)

? Pseudocloeon fastigatum Kopelke, 1980

? Pseudocloeon glaucum (Agnew, 1961) [South Africa]

? Baetis glaucus Agnew, 1961 (orig.)

? Baetis quintus Agnew, 1961 (syn.)

? Labiobaetis masai Lugo-Ortiz & McCafferty, 1977 (syn.)

? Labiobaetis nadineae Lugo-Ortiz & McCafferty, 1997 (syn.)

? Pseudocloeon masai Lugo-Ortiz, de Moor & Barber-James, 2000 (comb., syn)

? Pseudocloeon nadineae Lugo-Ortiz, de Moor & Barber-James, 2000 (comb., syn)

? Pseudocloeon quintum Lugo-Ortiz, de Moor & Barber-James, 2000 (comb., syn.)

? Pseudocloeon glaucum Lugo-Ortiz, McCafferty & Waltz, 1999 (comb.)

? Pseudocloeon grandiculum Kopelke, 1980

? Pseudocloeon insolitum (Kopelke, 1980)

? Baetis insolitus Kopelke, 1980 (orig.)

? Labiobaetis insolitum Lugo-Ortiz & McCafferty, 1997 (comb.)

? Pseudocloeon insolitum Lugo-Ortiz, McCafferty & Waltz 1999 (comb.)

? Pseudocloeon kalengoense (Kopelke, 1980)

? Baetis kalengoensis Kopelke, 1980 (orig.)

? Labiobaetis kalengoensis Lugo-Ortiz & McCafferty, 1997 (comb.)

? Pseudocloeon kalengoense Lugo-Ortiz, McCafferty & Waltz, 1999 (comb.)

? Pseudocloeon latum (Agnew, 1961) [South Africa]

? Baetis latus Agnew, 1961 (orig.)

? Labiobaetis aquacidus Lugo-Ortiz & McCafferty, 1997 (syn.)

? Pseudocloeon aquacidum Lugo-Ortiz, McCafferty & Waltz, 1999 (comb., syn.)

? Baetis sp. B. Harrison & Elsworth, 1958 (syn.)

? Baetis sp. 2. Agnew, 1961 (syn.)

? Pseudocloeon latum Lugo-Ortiz & de Moor, 2000 (comb.)

? Pseudocloeon mtone (Gillies, 1994)

? Baetis mtonis Gillies, 1994 (orig.)

? Labiobaetis mtonis Lugo-Ortiz & McCafferty, 1997 (comb.)

? Pseudocloeon mtone Lugo-Ortiz, McCafferty & Waltz, 1999 (comb.)

? Pseudocloeon piscis (Lugo-Ortiz & McCafferty, 1997) [South Africa]

? Labiobaetis piscis Lugo-Ortiz & McCafferty, 1997 (orig.)

? Pseudocloeon piscis Lugo-Ortiz, McCafferty & Waltz, 1999 (comb.)

? Pseudocloeon tenuicrinitum Kopelke, 1980

? Pseudocloeon vinosum Barnard, 1932 [South Africa]

? Pseudocloeon vinosum Barnard, 1932 (orig.)

? Baetis vinosus Gillies, 1994 (comb.)

? Pseudocloeon minutum Crass, 1947 (syn.)

? Baetis minutus Gillies, 1994 (comb., syn.)

? Labiobaetis vinosus Lugo-Ortiz & McCafferty, 1997 (comb.)

? Pseudocloeon vinosum Lugo-Ortiz, McCafferty & Waltz, 1999 (comb.)

33. Genus Pseudopannota Waltz & McCafferty, 1987

? Pseudopannota bergerardi Elouard & Hideux, 1990

? Pseudopannota bertrandi (Demoulin, 1967)

? Pseudocloeon bertrandi Demoulin, 1967 (orig.)

? Pseudopannota bertrandi Waltz & McCafferty, 1987 (comb.)

? Pseudopannota camillae Gattolliat (2002) [Madagascar]

? Pseudopannota maculosa (Crass, 1947) [South Africa]

? Pseudocloeon maculosum Crass, 1947 (orig.)

? Pseudopannota maculosa Elouard, Gillies & Wuillot, 1990 (comb.)

? Pseudopannota modesta Elouard & Gillies, 1990

? Pseudopannota muganinani Elouard & Gillies, 1990

? Pseudopannota sartorii Elouard & Hideux, 1990

? Pseudopannota vinckei (Demoulin, 1973) [Madagascar]

34. Genus Rheoptilum Gattolliat, 2001 [Madagascar]

? Rheoptilum arni Gattolliat, 2001 [Madagascar]

? Rheoptilum lokohensis Gattolliat, 2001 [Madagascar]

35. Genus Rhithrocloeon Gillies, 1985

? Rhithrocloeon indicator Gillies, 1985

? Rhithrocloeon permirum (Kopelke, 1980)

? Cloeon permirum Kopelke, 1980 (orig.)

? Rhithrocloeon permirum Gillies, 1985 (comb.)

36. Genus Scutoptilum Gattolliat, (2002) [Madagascar]

? Scutoptilum verrucosum Gattolliat (2002) [Madagascar]

37. Genus Securiops Jacobus, McCafferty and Gattolliat 2006

? Securiops macafertiorum (Lugo-Ortiz, 1996) [South Africa]

? Potamocloeon macafertiorum Lugo-Ortiz, 1996

? Potamocloeon dentatum Gattolliat, 2003 (syn.)

? Potamocloeon macafertiorum (Jacobus & McCafferty, 2005) (comb.)

? Secuirops macafertiorum Jacobus, McCafferty and Gattolliat 2006

? Securiops mandrare [Madagascar] Jacobus, McCafferty and Gattolliat 2006

? Securiops megapalpus Jacobus, McCafferty and Gattolliat 2006

? Securiops mutadens Jacobus, McCafferty and Gattolliat 2006

38. Genus Susua Lugo-Ortiz & McCafferty, 1998

? Susua niandanensis (Wuillot, 1993)

? Afroptilum niandanensis Wuillot, 1993 (orig.)

? Susua niandanensis Lugo-Ortiz & McCafferty, 1998 (comb.)

39. Genus Tanzaniops McCafferty & Barber-James 2005 [=Tanzaniella Gillies, 1991, n. hom., nec Tanzaniella Hoffman, 1977]

? Tanzaniops gorillorus (McCafferty, 2002)

? Tanzaniella gorillora McCafferty, 2002 (orig.)

? Tanzaniops gorillorus McCafferty& Barber-James, 2005 (comb.)

? Tanzaniops spinosus (Gillies, 1991)

? Tanzaniella spinosa Gillies, 1991 (orig)

? Tanzaniops spinosus McCafferty& Barber-James, 2005 (comb.)

40. Genus Thraulobaetodes Elouard & Hideux, 1991

? Thraulobaetodes cumminsorum Elouard & Hideux, 1991

41. Genus Xyrodromeus Lugo-Ortiz & McCafferty, 1997

? Xyrodromeus africanus Lugo-Ortiz &McCafferty, 1997

? Xyrodromeus latipalpus Gattolliat (2002) [Madagascar]

? Xyrodromeus namarona Lugo-Ortiz & McCafferty, 1997 [Madagascar]

? Xyrodromeus modestus Gattolliat (2002) [Madagascar]

? Xyrodromeus sartorii Gattolliat (2002) [Madagascar]

Suborder SETISURA

Superfamily HEPTAGENIOIDEA

Family HEPTAGENIIDAE Needham, 1901

1. Genus Afronurus Lestage, 1924

? Afronurus aethereus (Navàs, 1936)

? Ecdyonurus aethereus Navàs, 1936 (orig.)

? Afronurus aethereus Demoulin, 1965 (comb.)

? Afronurus barnardi Schoonbee, 1968 [South Africa]

? Afronurus elgonenis Puthz, 1971

? Afronurus gilliesi Corbet, 1962

? Afronurus harrisoni Barnard, 1932 [South Africa]

? Afronurus matitensis Sartori & Elouard, 1996

? Afronurus muehlenbergi Puthz, 1971

? Afronurus negi Cornert, 1960

? Afronurus oliffi Schoonbee, 1968 [South Africa]

? Afronurus peringueyi (Esben-Petersen, 1913)

? Ecdyurus peringueyi Esben-Petersen, 1913 (orig.)

? Afronurus peringueyi Lestage, 1924 (comb.)

? Afronurus pulcher Ulmer, 1930

? Adenophlebia collarti Navàs, 1930 (orig.)

? Afronurus collarti Demoulin, 1970 (comb.)

? Afronurus scotti Schoonbee, 1968 [South Africa]

? Afronurus subflavus Kopelke, 1980

? Afronurus ugandanus Kimmins, 1956 [South Africa]

2. Genus Compsoneuria Eaton, 1881

? Compsoneuria bequaerti (Navás,1930) [South Africa]

? Adenophlebia bequaerti Navás, 1930 (orig.)

? Adenophlebia eatoni Navás, 1931 (syn.)

? Adenophlebia inflexa Navás, 1932 (syn.)

? Notonurus cooperi Crass, 1947 (syn.)

? Notonurus bequaerti Demoulin, 1956 (comb.)

? Compsoneuriella bequaerti Gillies, 1963 (comb.)

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